Overview of Current Projects

Shark Systematics

Molecular systematics of sharks of the family Carcharhinidae (with Mahmood Shivji)

Shark Functional Morphology

Functional  morphology of the Greenland Shark (Somniosus microcephalus) splanchnocranium

Evolutionary and ecological correlates in egg case morphology of oviparous elasmobranchs (with Peter Bor and Hajimi Ishihara)

Shark Distribution

Zoogeography of Sharks: Dispersion versus Vicariance

Distribution and Diet of Greenland Sharks in the Saguenay Estuary (with Jeffrey Gallant and Frederic Milord)

Sharks of the Pacific Northwest

Distribution and strandings of White Sharks (Carcharodon carcharias) in British Columbia and Alaska waters.

Population dynamics of Bluntnose Sixgill Sharks (Hexanchus griseus) at Hornby Island, British Columbia

Identity of a possible second species of deep-sea catshark (Apristurus sp.) off British Columbia, Canada

White Sharks

Diet, predatory behavior, and feeding ecology of White Sharks: a critical review

Predator-prey relationships between White Sharks and Cape Fur Seals (Arctocephalus pusillus pusillus) at False Bay, South Africa

Application of Geographic Profiling to the study of White Shark predatory behaviour at Seal Island, South Africa (with Kim Rossmo and Neil Hammerschlag)

Short- and long-term identification of individual White Sharks at Seal Island, South Africa, based on pigmentation and scarification patterns (with Kathryn Hodgson, Gennadyi Gurman, Neil Hammerschlag, and Chris Fallows)

Attack on seabirds by White Sharks at Seal Island, South Africa:  mistaken identity, investigation, or play? (with Neil Hammerschlag, Kathryn Hodgson, and Chris Fallows)

Deep-Sea Sharks

Bioluminescence and squamation of deep-sea squaloids (families Etmopteridae and Dalatiidae)

Biology of deep-sea sharks: a review (with Jason Treberg)

Muscle enzyme activity in the deep sea Black Dogfish (Centroscyllium fabricii) compared with that in its shallow-living relative the Spiny Dogfish (Squalus acanthias) (with Jason Treberg and William Driedzic)

Filter-Feeding Elasmobranchs

Feeding ecology of the Manta Ray (Manta birostris) in Yap, western Caroline Islands

Functional significance of pigmentation pattern in the Whale Shark (Rhincodon typus) (with Steven Wilson)

Shark Social Behavior

Agonistic displays in tropical reef sharks (family Carcharhinidae) of the Indo-West Pacific region

Agonistic displays in two species of squalean shark (Hexanchus griseus, Squalus acanthias)

Natural Mortality of Sharks

Mortality of Puffadder Shysharks (Haploblepharus edwardsii) due to Cape Fur Seals (Arctocephalus pusillus pusillus) and Kelp Gulls (Larus dominicanis) at Seal Island, South Africa.

Shark Attacks

Seasonal attacks by White Sharks on surfers off California and Oregon (with Ralph Collier and Dan Warner)

Identification Guides

Guide to Sharks of the World: Identification, Distribution, Life History, Utilisation, and Conservation (book)

Field Guide to the Sharks of the Pacific Coast of North America (with Alessandro De Maddalena)